Most definitely cooler in the air. I have found that I am way to rich by way of ground tuning once they get airborn. But that works out well and I would not recommend tuning for only flight temps. A DLE 55 was the first of my Tele experiences, on the ground, properly tuned, it was running 230°, but as soon as the wheels left the ground it came down and ran pretty consistently around 190°. With the exception of a chopped throttle, the longer at idle the higher the temp went, back on throttle and down it came. A really cool thing to watch especially with rpm, airspeed, and throttle position. I record mine and watch on the ground, after the fact, watching in flight is like texting while driving. Don't do it!!