RPM and cooling are the big differences. We don't try to ring out every RPM we can get from an engine and a planes cooling system is way better. RPM can be controlled with prop selection. When tuned we run up a plane engine to the max RPM we can get then back off 4 or 5 hundred RPM rich. Between that and the cooling, our plane engines last for years without ever requiring a rebuild. Only time I touch my needle is during big weather changes or when I change prop sizes. The fuel and oil type and content can effect the life of a plane engine but that's another topic.