Hydro Junkie

Nope, neither one. Neither of them saw combat in two conflicts. The P-38 was retired after VJ Day while the F-82 wasn't ready in time for WWII and barely finished the Korean War. I'll give a partial run-down of the clues and see if it helps:
1) These two aircraft shared a distinction from all other aircraft
2) While both types had great combat records, one was the first and the other the last
3) The two aircraft had maiden flights approximately four years apart The first flew the first time 25 November 1940, the last April 1945
4) The two aircraft made their first combat flights approximately nine years apart The first flew a photo/recon mission 17 September 1941, the last first flew 3 July 1950
5) The two aircraft were involved in events that led to the loss of the first but not the second
6) Only the aircraft that was lost had more than one person on board. The first had a crew or two or three, the last had only a pilot
7) The "last" aircraft was performing a function it was never designed to perform
8) The "last" aircraft was replaced soon after by new, faster aircraft that were themselves in service for well over 30 years The last was replaced by the A-4 Skyhawk, A-6 Intruder and A-7 Corsair II
9) The last of the "first" was retired by the air forces of the building nation after only eight years of service The last one was retired in 1950
10) Both the "first" and the "last" had a disadvantage in their similar events, the "first" wasn't able to overcome this while the "last" did Both were outperformed in these events
11) The aircraft involved all used different engines
12) All aircraft involved were armed with either machine guns or cannons, no missiles
13) The "first" and "last" flyers spoke the same language, more or less, while everyone else involved spoke their own Only the flight crews of the first and last spoke a form of English
14) The events started by the "first"(resulting in it's crash landing afterward) and ended with the "last" covered three major conflicts involving the nation that manufactured the "last"
15) The "first" used a radical construction(for the time it was designed) that the powers that be didn't really trust or want to buy at first while the "last" was designed and used to do the same tasks as three well known, front line, proven combat aircraft that it replaced The first used wood in a previously untried way, the last replaced the SB2-C Helldiver, TBF/TBM Avenger and F4U Corsair
16) The "first" was this country's first aircraft to top the 400 MPH mark while the "last" was the last not to make 400 for it's country,even though it had more power than many that could
17) Both aircraft were designed during the same conflict, the "first" participated while the "last" waited for the next where it did not fly in defense of it's own country but to protect troops from it
Let's see if that gives it away